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After the defeat of Mechagodzilla,[b] Kong establishes his territory in the recently discovered Hollow Earth. While on Earth's surface, Godzilla continues to maintain order between humanity and giant monsters, known as "Titans" — defeating Scylla in Rome and resting in the Colosseum afterwards. A Monarch observation outpost stationed in the Hollow Earth picks up an unidentified signal. On the surface, the signal causes Jia, the last member of the Iwi tribe, to experience hallucinations and flashbacks, causing her and her adopted mother, Dr. Ilene Andrews, to worry. Also sensing the signal, Godzilla leaves Rome, attacks a nuclear plant in France to absorb the radiation, and travels to the Arctic; Monarch believes Godzilla is strengthening himself for an oncoming threat.
When a sinkhole opens near his home, Kong discovers an uncharted region and a tribe of giant apes — including an adolescent, Suko. After an initial confrontation, Kong forces Suko to lead him to the tribe's lair and the two slowly bond on their journey. The tribe's tyrannical leader, the Skar King, battles and injures Kong with the aid of an ancient ice-powered Titan, Shimo, that Skar King controls with pain using a crystal.
Andrews and Jia, alongside Titan veterinarian Trapper and conspiracy podcaster Bernie Hayes, travel to the Hollow Earth to locate the source of the signal and find the Monarch outpost destroyed. As the group ventures forward, they discover an uncharted subterranean section that houses the remaining members of the Iwi tribe, a group of people who communicate telepathically among each other and from whom Jia is descended. Inspecting some ruins, they surmise that the signal was a telepathic distress call sent by the Iwi.
As Andrews observes Jia socializing with the Iwi, she voices her fears to Trapper that Jia may choose to stay with her people and Andrews would have to accept it. Inside a temple, Andrews uncovers a prophecy: long ago, Skar King attempted to take over the surface world, and waged war against Godzilla's species in the process; Godzilla succeeded in trapping Skar King and his tribe deep within the Hollow Earth. The prophecy also states that Jia would be the key to reawakening Mothra. Kong, sensing Jia, locates the temple and is fitted by Trapper with a prototypical prosthetic battle-glove to treat his wounds.
Godzilla defeats the monster Tiamat in the Arctic in order to absorb the radiation from its lair. Hoping to lure Godzilla to the Hollow Earth to assist him in battle, Kong confronts him in Cairo. An empowered Godzilla gains the upper hand, but the reawakened Mothra intervenes to convince him to aid Kong. Godzilla, Kong, and Mothra battle with Skar King, his tribe, and Shimo in the Hollow Earth. Their fight carries them into Rio de Janeiro, where Shimo begins to induce a second Ice Age at Skar King's command. The factions are evenly matched until Suko destroys the crystal used to control Shimo. Finally liberated, Shimo freezes Skar King and Kong shatters his frozen body.
After the battle, Godzilla return